Unearthed teeth reveal human diversity in China during Middle Pleistocene

Gaby Clark
scientific editor

Andrew Zinin
lead editor

The Centro Nacional de Investigación sobre la Evolución Humana (CENIEH) participated in a study, published in the , on human teeth unearthed at the Hualongdong site in Anhui Province, China. The findings offer fresh insights into hominin diversity in Asia during the late Middle Pleistocene. Analysis of these fossil teeth reveals an unusual combination of primitive and modern traits that challenges established classifications of Homo evolution on the Asian continent.
Led by Professor Wu Xiujie, director of the Hualongdong excavations, the research is the result of a long-standing collaboration between scientists from the Dental Anthropology Group at CENIEH—MarÃa Martinón-Torres, Director of CENIEH and corresponding author of the paper, and José MarÃa Bermúdez de Castro, ad Honorem researcher at CENIEH— and scientists from the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing.
The Hualongdong fossil assemblage includes a nearly complete cranium with 14 teeth preserved in situ, a partial maxilla, several isolated teeth, femoral fragments, and other cranial remains. Through a comparative analysis of 21 dental elements, the researchers identified a morphological pattern that combines archaic features—such as robust dental roots typical of the Middle Pleistocene—with more modern traits, including a reduced third molar commonly found in Homo sapiens and other Late Pleistocene hominins.
However, the Hualongdong fossils lack the dental features characteristic of Neanderthals, suggesting a distinct population. This blend of traits could be the result of various evolutionary processes, ranging from genetic admixture with more primitive species like Homo erectus to the possibility that these individuals represent a lineage related to Homo sapiens, yet distinct from both Neanderthals and Denisovans.
An evolutionary mosaic
The findings at Hualongdong reinforce the idea that traits associated with the emergence of Homo sapiens were already present in Asia at least 300,000 years ago. While the facial region of these hominins exhibits modern-like features, certain dental and mandibular characteristics remain distinctly archaic.
"It's a mosaic of primitive and derived traits never seen before—almost as if the evolutionary clock were ticking at different speeds in different parts of the body," explains MarÃa Martinón-Torres. According to José MarÃa Bermúdez de Castro, "the Hualongdong discovery reminds us that human evolution was neither linear nor uniform, and that Asia hosted multiple evolutionary experiments with unique anatomical outcomes."
These results further support the view that human evolution in Asia during the Pleistocene was more complex and diverse than previously thought—particularly given that a at the same site had already identified a juvenile individual exhibiting a similar combination of modern and archaic traits.
The Hualongdong site now joins other key fossil localities, such as Panxian Dadong and Jinniushan, which may offer further insights into the evolutionary pathways that led to modern humans.
More information: Xiujie Wu et al, The hominin teeth from the late Middle Pleistocene Hualongdong site, China, Journal of Human Evolution (2025).
Journal information: Journal of Human Evolution